• University of California, San Diego, La Jolla, CA, 1976-1982, PhD in Biology, 1982, Dissertation: Isolation and characterization of DI particle resistant mutants of vesicular stomatitis virus.
   
• Brown University, Providence, RI, 1971-1975, ScB in Biochemistry, Summa cum laude with honors, 1975.

Positions Held:

• Professor, Department of Biomedical Sciences and the Heritage College of Osteopathic Medicine, 2004-present.
   
• Associate Professor, Department of Biomedical Sciences and the Heritage College of Osteopathic Medicine, Ohio University, 1999-2004.
   
• Assistant Professor, Department of Biomedical Sciences and the Heritage College of Osteopathic Medicine, Ohio University, 1998-1999.
   
• Assistant Professor, Department of Biological Sciences and the Molecular and Cellular Biology Program, Ohio University, 1993-1998.
   
• Assistant Professor, Department of Biochemistry, University of Nevada, Reno, 1990-1993.
   
• Postdoctoral research in insect molecular biology, University of Washington, Zoology Department, under Dr. Lynn Riddiford, 1984-1990.
   
• Postdoctoral research in plant molecular biology, University of Washington, Biochemistry Department, under Dr. Milton Gordon, 1983.
   
• Graduate research in virology, University of California, San Diego, Biology Department, under Dr. John Holland, 1977-1982.
   
• Research technician in yeast molecular biology, University of Washington, Genetics Department, under Dr. Benjamin Hall, 1975-1976.