Hieronymus, T.L., and L.M. Witmer (in review) Adaptation, exaptation, and convergence in rhinocerotid skin evolution. Submitted to Proceedings of the National Academy of Sciences. 

Hieronymus, T.L. (in press) Comparative anatomy and physiology of chemical senses in aquatic birds. In: Thewissen, J.G.M., and S. Nummela (eds), Sensory Evolution on the Threshold, Adaptations in Secondarily Aquatic Vertebrates. University of California Press. 

Thewissen, J.G.M., and T.L. Hieronymus (in press) Evolutionary relationships of aquatic birds. In: Thewissen, J.G.M., and S. Nummela (eds), Sensory Evolution on the Threshold, Adaptations in Secondarily Aquatic Vertebrates. University of California Press. 

Hieronymus, T.L., L.M. Witmer, and R. Ridgely (2006) The structure of white rhinoceros (Ceratotherium simum) horn investigated by X-ray computed tomography and histology, with implications for growth and external form. Journal of Morphology 267:1172–1176.  (PDF)

Hieronymus, T.L. (2006) Quantitative microanatomy of jaw muscle attachment in extant diapsids. Journal of Morphology 267:954–967. (PDF) 

Organ, C.L., J.B. Cooley, and T.L. Hieronymus (2003) A non-invasive quarry mapping system. Palaios 18(1): 74–77. (PDF)

Published Abstracts

Hieronymus, T.L., and L.M. Witmer. (2007) Skinning dinosaurs: bony correlates and patterns of cephalic evolution in Archosauria. Journal of Vertebrate Paleontology 27(3, suppl.): 89A.

Hieronymus, T.L., and L.M. Witmer. (2007) Turtle beaks, bird beaks, croc beaks? Parallel evolution of rhamphothecae in Sauropsida. Journal of Morphology 268(12):1083-1084.

Hieronymus, T.L., and L.M. Witmer. (2007) How dinosaurs build beaks: Homology between avian rhampotheca and diapsid facial scales. Integrative and Comparative Biology 45(6, suppl.): e60. 

Hieronymus, T.L., L.M. Witmer, D. Tanke, and P.J. Currie. (2006) Horn morphology of Pachyrhinosaurus and horn evolution in centrosaurine dinosaurs. Journal of Vertebrate Paleontology 26(3, suppl): 75A. 

Roberts, E.M., P.M. O'Connor, T.L. Hieronymus, S. Ngasala, M.D. Gottfried, and N.J. Stevens. (2006) Stratigraphy, paleoenvironments, and taphonomy of a new Cretaceous Vertebrate fauna from the East African rift, southwestern Tanzania; p. 175 in Barrett, P. M., and S. E. Evans (eds), Ninth International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings. 

Hieronymus, T.L., and L.M. Witmer (2006) From armor to anchor: Character evolution in the skin and horns of rhinos. Integrative and Comparative Biology 45(6): 1012. 

Hieronymus, T.L., and L.M. Witmer (2005) Revised horn reconstructions for extinct rhinocerotid taxa: Separating the contributions of dermis and epidermis to horn rugosities. Journal of Vertebrate Paleontology 25(3, suppl.): 69A. 

Witmer, L., R. Ridgely, P. O'Connor, C. Holliday, T. Hieronymus, J. Daniel, E. Rasmusson, J. Tickhill, and A. Clifford (2005) The cephalic anatomy of the Florida manatee (Trichechus manatus) as revealed by virtual dissection with computed tomography (CT), differential-contrast dual-vascular injection, and 3D visualization. Evolution of Aquatic Tetrapods, Fourth Triannual Conference. 

Hieronymus, T.L., and L.M. Witmer (2004) Cranial rugosity and dinosaur "horns:" Rhino and giraffe as model systems for skin reconstruction in fossil taxa. Journal of Vertebrate Paleontology 24(3, suppl.): 70A. 

Hieronymus, T.L., and L.M. Witmer (2004) Rhinoceros horn attachment: anatomy and histology of a dermally influenced bone rugosity. Journal of Morphology 260(3): 298. 

Hieronymus, T.L., and L.M Witmer (2004) Dermal enthesis: anatomy and histology of rhinoceros horn attachment. Integrative and Comparative Biology 43(6): 874. 

Hieronymus, T.L., and L.M. Witmer (2003) Rhinoceros horn attachment: anatomy and histology. Journal of Vertebrate Paleontology 23(3, suppl.): 61A. 

Hieronymus, T.L. (2003) Microanatomical correlates of muscle attachment and their implications for muscular reconstruction. Integrative and Comparative Biology 42(6): 1243. 

Hieronymus, T.L. (2002) Microanatomical and histological correlates of muscle attachment and implications for muscular reconstruction. Journal of Vertebrate Paleontology 22(3, suppl.): 64A. 

Hieronymus, T.L. (2001) Sharpey’s fiber pits on the periosteal surface of a pachycephalosaurid dome. Journal of Vertebrate Paleontology 21(3, suppl.): 61A. 

Cooley, J.B., T.L. Hieronymus, and C.L. Organ (2001) A non-invasive floating grid mapping system. PaleoBios 21(2, suppl.): 43.